VMware vSAN all versions.
The issue is in the upstream physical network layer.
Because the state is held upstream, rebooting the ESXi hosts fails to resolve the issue, as the reboot does not force the physical switch to clear the specific network state associated with the uplinks.
The network layer needs to be investigated.
Refreshing the physical link (Link-Down/Link-Up) resolves issues where the upstream network has cached a bad state or encountered a buffer/protocol hang that is not sensitive to host-side software restarts.